The Leucine zipper-EF-hand-containing transmembrane protein1 (LETM1), located at the inner mitochondrial membrane, has been indicated to fuction as the K+/H+ antiporter, and LETM1-depleted mitochondria accumulate high K+ levels inside their matrix. Moreover, it has been proposed that the LETM1-dependent Ca2+ efflux is modulated by mitochondrial Na+ homeostasis.
References:
Nowikovsky K, Froschauer EM, Zsurka G, Samaj J, Reipert S, Kolisek M, Wiesenberger G, Schweyen RJ. The LETM1/YOL027 gene family encodes a factor of the mitochondrial K+ homeostasis with a potential role in the Wolf-Hirschhorn syndrome. J Biol Chem. 2004 Jul 16;279(29):30307-15. doi: 10.1074/jbc.M403607200. Epub 2004 May 11. PMID: 15138253.
De Marchi U, Santo-Domingo J, Castelbou C, Sekler I, Wiederkehr A, Demaurex N. NCLX protein, but not LETM1, mediates mitochondrial Ca2+ extrusion, thereby limiting Ca2+-induced NAD(P)H production and modulating matrix redox state. J Biol Chem. 2014 Jul 18;289(29):20377-85. doi: 10.1074/jbc.M113.540898. Epub 2014 Jun 4. PMID: 24898248.
Austin S, Tavakoli M, Pfeiffer C, Seifert J, Mattarei A, De Stefani D, Zoratti M, Nowikovsky K. LETM1-Mediated K+ and Na+
Homeostasis Regulates Mitochondrial Ca2+ Efflux. Front Physiol. 2017 Nov 17;8:839. doi: 10.3389/fphys.2017.00839. PMID: 29204122.